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Mitrophan Znosko-Borovsky was born into the family of a parish priest on August 17 (August 4 O.S.), 1909, in the city of Brest-Litovsk (at that time a part of the Russian Empire).
Mitrophan was ordained to priesthood on June 12th, 1936, shortly before he married his matushka Alexandra Semenovna Tsibrouk.